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t temporal patterning predicts stimulus-dependent synchrony in nearby cells sharing input with similar spectrotemporal structure, which in turn can sharpen the downstream readout of sound direction. This work shows how stimulus-dependent spike timing can affect the downstream coding of sound location by firing rate, a mechanism that can be generalized to sensory neurons sensitive to the temporal structure of the stimulus. Copyright © 2020 Beckert et al.Altered synaptic function is thought to play a role in ma