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These fundamental processes of symmetry breaking, axiation, patterning, and morphogenesis have been extensively studied in developmental biology but less so at the subcellular level. This review will focus on developmental processes that give eukaryotic cells their complex structures, with a focus on cytoskeletal organization in free-living cells, ciliates in particular, in which these processes are most readily apparent.Philip Donoghue introduces the fossil record of cells.Medina and Buchler provide an introduction to chytrid fungi, an
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