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tial connectivity. © 2020 The Authors. Ecology and Evolution published by John Wiley Sons Ltd.There has been much recent research interest in the existence of a major axis of life-history variation along a fast-slow continuum within almost all major taxonomic groups. Eco-evolutionary models of density-dependent selection provide a general explanation for such observations of interspecific variation in the "pace of life." One issue, however, is that some large-bodied long-lived "slow" species (e.g., trees and lar