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In contrast, leaf traits were affected by the variation in phenology within crowns-nitrogen content and photosynthetic capacity were greater in the leaves of the warmed branches than in the control branches, but they did not differ between the leaves of warmed trees and control trees. Thus, branch warming altered the distribution of nitrogen between warmed and unwarmed branches as warmed branches developed faster, resulting in intracrown variation in leaf photosynthetic traits.Since 2002, Alaria (A.) alata mesocercariae (AM) have been fou
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