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mariesii in canopy gaps, and vice versa in the forest understory; and (2) at high elevation, the surplus production rate of A. mariesii is greater than that of A. veitchii in both forest understory and canopy gaps. The results obtained in this study were consistent with our two stated hypotheses. In addition, at the low elevation site, the rate of the growth in height of A. veitchii in canopy gaps was greater than that of A. mariesii, indicating that A. veitchii can dominate after disturbance at low elevations. The findings of this study