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At cell level, all OsEULs are located in the nucleus whereas OsEULD1b and OsEULD2 also locate to the cytoplasm. This paper contributes to the functional characterization of the EULs and provides insight in the biological importance of this family of proteins for rice. Copyright © 2020 Lambin, Demirel Asci, Dubiel, Tsaneva, Verbeke, Wytynck, De Zaeytijd, Smagghe, Subramanyam and Van Damme.Epigenetics refers to the mode of inheritance independent of mutational changes in the DNA. Early evidence has revealed methylation, acetylation,