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These outcomes indicate that YcjR catalyzes the isomerization of 3-keto-d-glucosides via proton abstraction at C4 by Glu-146 to form a cis-enediolate intermediate that is afterwards protonated regarding the reverse face by Glu-240 to generate the corresponding 3-keto-d-guloside. This conclusion is sustained by docking regarding the cis-enediolate intermediate into the energetic web site of YcjR in line with the understood binding orientation of d-fructose and d-psicose when you look at the active web sit
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